Everything about The Cougar totally explained
The
cougar (
Puma concolor), also
Puma,
mountain lion, or
panther, is a
mammal of the
Felidae family, native to the
Americas. This large, solitary cat has the greatest
range of any wild terrestrial mammal in the Western Hemisphere,
"Cougar" is borrowed from the
Portuguese çuçuarana, via
French; the term was originally derived from the
Tupi language. A current form in Brazil is
suçuarana. "Puma" comes, via
Spanish, from the
Quechua language of Peru.
Taxonomy and evolution
The cougar is the largest of the small cats. It is placed in the subfamily
Felinae, although its bulk characteristics are similar to those of the
big cats in the subfamily
Pantherinae. as cats are poorly represented in the
fossil record, North American felids then invaded South America 3 Ma ago as part of the
Great American Interchange, following formation of the
Isthmus of Panama. The cougar was originally thought to belong in
Felis, the genus which includes the
domestic cat, but it's now placed in
Puma along with the
jaguarundi, a cat just a little more than a tenth its weight.
Studies have indicated that the cougar and jaguarundi are most closely related to the modern
cheetah of Africa and western Asia, but the relationship is unresolved. It has been suggested that the cheetah lineage diverged from the
Puma lineage in the Americas (see
American cheetah) and migrated back to Asia and Africa, The outline of
small feline migration to the Americas is thus unclear.
Recent studies have demonstrated a high level of genetic similarity among the North American cougar populations, suggesting that they're all fairly recent descendants of a small ancestral group. Culver
et al. suggest that the original North American population of
Puma concolor was
extirpated during the
Pleistocene extinctions some 10,000 years ago, when other large mammals such as
Smilodon also disappeared. North America was then repopulated by a group of South American cougars.
Argentine puma : includes the previous subspecies and synonyms hudsonii and puma (Marcelli, 1922);
; Costa Rican Cougar
Eastern South American cougar : includes the previous subspecies and synonyms acrocodia, borbensis, capricornensis, concolor (Pelzeln, 1883), greeni and nigra;
; North American Cougar : includes the previous subspecies and synonyms arundivaga, aztecus, browni, californica, coryi, floridana, hippolestes, improcera, kaibabensis, mayensis, missoulensis, olympus, oregonensis, schorgeri, stanleyana, vancouverensis and youngi;
Northern South American cougar : includes the previous subspecies and synonyms bangsi, incarum, osgoodi, soasoaranna, soderstromii, sucuacuara and wavula;
; Southern South American puma : includes the previous subspecies and synonyms araucanus, concolor (Gay, 1847), patagonica, pearsoni and puma (Trouessart, 1904)
The status of the Florida panther, here collapsed into the North American cougar, remains uncertain. It is still regularly listed as subspecies Puma concolor coryi in research works, including those directly concerned with its conservation. Culver et al. themselves noted microsatellite variation in the Florida panther, possibly due to inbreeding;
Biology and behavior
Physical characteristics
Cougars are slender and agile cats. Adults stand about 60 to 80 cm (2.0 to 2.7 ft) tall at the shoulders. The length of adult males is around 2.4 m (8 ft) long nose to tail, with overall ranges between 1.5 and 2.75 meters (5 and 9 ft) nose to tail suggested for the species in general. Males have an average weight of about 53 to 72 kilograms (115 to 160 pounds). In rare cases, some may reach over 120 kg (260 lb). Female average weight is between 34 and 48 kg (75 and 105 lb). Cougar size is smallest close to the
equator, and larger towards the
poles.
The head of the cat is round and the ears erect. Its powerful forequarters, neck, and jaw serve to grasp and hold large prey. It has five retractable claws on its forepaws (one a
dewclaw) and four on its hind paws. The larger front feet and claws are adaptations to clutching prey.
Cougars can be as large as jaguars, but are less muscled and powerful; where their ranges overlap, the cougar tends to be smaller than average. The cougar is on average heavier than the leopard. Despite its size, it isn't typically classified among the "
big cats," as it can't roar, lacking the specialized
larynx and
hyoid apparatus of
Panthera. Like domestic cats, cougars vocalize low-pitched hisses, growls, and purrs, as well as chirps and whistles. They are well known for their screams, referenced in some of its common names, although these may often be the misinterpreted calls of other animals.
Cougar coloring is plain (hence the
Latin concolor) but can vary greatly between individuals and even between siblings. The coat is typically tawny, but ranges to silvery-grey or reddish, with lighter patches on the under body including the jaws, chin, and throat. Infants are spotted and born with blue eyes and rings on their tails; The term "
black panther" is used colloquially to refer to melanistic individuals of other species, particularly jaguars and leopards.
Cougars have large paws and proportionally the largest hind legs in the cat family. Horizontal jumping capability is suggested anywhere from 6 to 12 m (20 to 40 ft). The cougar can run as fast as 55 km/h (35 mph), but is best adapted for short, powerful sprints rather than long chases. It is adept at climbing, which allows it to evade
canine competitors. Although it isn't strongly associated with water, it can swim.
Hunting and diet
A successful
generalist predator, the cougar will eat any animal it can catch, from
insects to large
ungulates. Like all cats, it's an
obligate carnivore, feeding only on meat. Its most important prey species are various deer species, particularly in North America;
mule deer,
white-tailed deer,
elk, and even the large
moose are taken by the cat. Other species such as Bighorn Sheep, horses, and domestic livestock such as cattle and sheep are also primary food bases in many areas. A survey of North America research found 68% of prey items were ungulates, especially deer. Only the Florida Panther showed variation, often preferring feral
hogs and
armadillos. Another study on winter kills (November–April) in
Alberta showed that ungulates accounted for greater than 99% of the cougar diet. Learned, individual prey recognition was observed, as some cougars rarely killed
bighorn sheep, while others relied heavily on the species. The cougar also has conflicts with species such as
American Black Bear and
Brown Bear, as in many cases the bears will displace the cougar and consume their kills, forcing the cougar to kill larger than normal numbers of prey animals. Cougars may themselves displace smaller carnivores for their own kills, such as
Coyotes.
In the Central and South American cougar range, the ratio of deer in the diet declines. Small to mid-size mammals are preferred, including large rodents such as the
capybara. Ungulates accounted for only 35% of prey items in one survey, approximately half that of North America. Competition with the larger jaguar has been suggested for the decline in the size of prey items.
Reproduction and lifecycle
Females reach sexual maturity between one-and-a-half and three years of age. They typically average one
litter every two to three years throughout their reproductive life; the period can be as short as one year. Copulation is brief but frequent.
Only females are involved in parenting. Female cougars are fiercely protective of their kittens, and have been seen to successfully fight off animals as large as grizzly bears in their defense. Litter size is between one and six kittens, typically two or three. Caves and other alcoves that offer protection are used as litter dens. Born blind, kittens are completely dependent on their mother at first, and begin to be weaned at around three months of age. As they grow, they begin to go out on forays with their mother, first visiting kill sites, and after six months beginning to hunt small prey on their own.
Life expectancy in the wild is reported at between 8 to 13 years, and probably averages 8 to 10; a female of at least 18 years was reported killed by hunters on
Vancouver Island.
Social structure and home range
Like almost all cats, the cougar is a solitary animal. Only mothers and kittens live in groups, with adults meeting only to mate. It is secretive and
crepuscular, being most active around dawn and dusk.
Estimates of territory sizes vary greatly.
Canadian Geographic reports large male territories of 150 to 1000
square kilometers (58 to 386
sq mi) with female ranges half the size. Male ranges may include or overlap with those of females but, at least where studied, not with those of other males, which serves to reduce conflict between cougars. Ranges of females may overlap slightly with each other. Scrape marks,
urine, and
feces are used to mark territory and attract mates. Males may scrape together a small pile of leaves and grasses and then urinate on it as a way of marking territory.
Ecology
Distribution and habitat
The cougar has the largest range of any wild land animal in the Americas. Its range spans 110 degrees of
latitude, from northern
Yukon in Canada to the southern
Andes. It is one of only three cat species, along with the
bobcat and
Canadian lynx, native to Canada. DNA evidence has suggested its presence in eastern North America, while a consolidated map of cougar sightings shows numerous reports, from the mid-western
Great Plains through to
Eastern Canada.
Genetic analysis of DNA from a cougar sighting in Wisconsin in 2008 indicated that a cougar was in Wisconsin and that it wasn't captive. It is speculated that the cougar migrated from a native population in the Black Hills of South Dakota; however, the genetic analysis couldn't affirm that hypothesis. It is also uncertain whether there are other, perhaps breeding, cougars. However, a second sighting was reported and tracks were documented in a nearby Wisconsin community. Unfortunately, a genetic analysis couldn't be done and a determination couldn't be made. This cougar later made its way south into the northern
Chicago suburb of
Wilmette. On
April 14,
2008, a cougar triggered a flurry of reports before being cornered and killed in the Chicago neighborhood of
Roscoe Village while officers tried to contain it. The cougar was the first sighted in the city limits of Chicago since the city was founded in 1833. The only unequivocally known eastern population is the
Florida panther, which is critically endangered.
South of the
Rio Grande, the
International Union for the Conservation of Nature and Natural Resources (IUCN) lists the cat in every
Central and
South American country except
Costa Rica and
Panama.
The cougar's total breeding population is estimated at less than 50,000 by the IUCN, with a declining trend. California has actively sought to protect the cat and a similar number of cougars has been suggested, between 4,000 and 6,000.
Ecological role
Aside from humans, no species preys upon mature cougars in the wild. The cat is not, however, the
apex predator throughout much of its range. In its northern range, the cougar interacts with other powerful predators such as the
brown bear and gray wolf (although a lone wolf poses little threat to an adult cougar). In the south, the cougar must compete with the larger jaguar. In Florida it encounters the
American alligator.
The
World Conservation Union (IUCN) currently lists the cougar as a "
near threatened" species. It has shifted the cougar's status from "least concern," while leaving open the possibility that it may be raised to "
vulnerable" when greater data on the cat's distribution becomes available. rendering illegal international trade in specimens or parts.
East of the
Mississippi, the only unequivocally known cougar population is the
Florida panther. The
U.S. Fish and Wildlife Service recognizes both an Eastern cougar and the Florida panther, affording protection under the
Endangered Species Act. Certain taxonomic authorities have collapsed both designations into the
North American Cougar, with Eastern or Florida subspecies not recognized,
The cougar is also protected across much of the rest of its range.
As of 1996, cougar hunting was prohibited in
Argentina,
Brazil,
Bolivia,
Chile,
Colombia,
Costa Rica,
French Guiana,
Guatemala,
Honduras,
Nicaragua,
Panama,
Paraguay,
Suriname,
Venezuela, and
Uruguay. (Costa Rica and Panama are not listed as current range countries by the IUCN.) The cat had no reported legal protection in
Ecuador,
El Salvador, and
Guyana.
Attacks on humans
Due to the growth of
urban areas, cougar ranges increasingly overlap with areas inhabited by
humans. Attacks on humans are rare, as cougar prey recognition is a learned behavior and they don't generally recognize humans as prey. Attacks on people, livestock, and pets may occur when the cat
habituates to humans. There have been 108 confirmed attacks on humans with twenty fatalities in North America since 1890, fifty of the incidents having occurred since 1991. The heavily populated state of
California has seen a dozen attacks since 1986 (after just three from 1890 to 1985), including three fatalities.
When the cougar does attack, it usually employs its characteristic neck bite, attempting to position its teeth between the
vertebrae and into the
spinal cord. Neck, head, and spinal injuries are common and sometimes fatal. The sky and thunder god of the Inca,
Viracocha, has been associated with the animal.
In North America, mythological descriptions of the cougar have appeared in the stories of the
Hotcâk language ("Ho-Chunk" or "Winnebago") of
Wisconsin and
Illinois and the
Cheyenne, amongst others. To the
Apache and
Walapai of
Arizona, the wail of the Cougar was harbinger of death.
The cougar continues to be a symbol of strength and stealth. From
combat helicopters, motor vehicles (see
Ford/Mercury Cougar and
Ford Puma) to
athletic shoes, both "Cougar" and "Puma" are widely used as
brand names. Various sports teams have also adopted the names, including the
Argentina national rugby union team as well as US universities,
Brigham Young University and
Washington State University. Many places, such as
Cougar Mountain, are also named after their association with cougars.
Further Information
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